I have a theory, just a small one, about Connor in the Tyranny of King Washington DLC. Spoilers below the break. 

Connor’s eyes turned blue after the last trip to the spirit world. Wolf cubs’ eyes are blue when they are born, and turn to amber or brown or even green as they mature. He is just like a wolf cub. And I’m sure if he had been allowed to continue this way, his eyes would have changed to amber as well.

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